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ganglionic structures and the funneling of efferent projections to the ventrolateral thalamus and then to the motor cortex, current evidence indicates an organization into several parallel basal ganglionic cortical circuits Although these circuits run parallel to the premotor or prepyramidal pathway, they remain separate from it anatomically and physiologically At least ve such circuits have been described, each projecting to a different portion of the frontal lobe: (1) the prototypical motor circuit, converging on the premotor cortex; (2) the oculomotor circuit, on the frontal eye elds; two prefrontal circuits, (3) one on the dorsolateral prefrontal and (4) another on the lateral orbitofrontal cortex; and (5) a limbic circuit that projects to the anterior cingulate and medial orbitofrontal cortex An additional and essential feature of basal ganglionic structure, also relatively recently appreciated, is the nonequivalence of all parts of the striatum Particular cell types and zones of cells within this structure appear to mediate different aspects of motor control and to utilize speci c neurochemical transmitters, as detailed below under Pharmacologic Considerations (see Albin et al and DeLong) This specialization has taken on further importance with the observation that one or another cell type is found to be destroyed preferentially in degenerative diseases such as Huntington chorea The most important basal ganglionic connections and circuitry are indicated in Figs 4-1, 4-2, and 4-3 Conventionally it has been taught that the striatum, mainly the putamen, is the receptive part of the basal ganglia, receiving topographically organized bers from all parts of the cerebral cortex and from the pars compacta (pigmented neurons) of the substantia nigra, and that the output nuclei of the basal ganglia consist of the medial (internal) pallidum and the pars reticulata (nonpigmented portion) of the substantia nigra (Fig 4-3) It has been proposed on the basis of physiologic, lesional, and pharmacologic studies that there are two main efferent projections from the putamen; but these models, being quite incomplete, should be viewed as provisional Nonetheless, there are reasons to conceptualize a direct efferent system from the putamen onto the medial (internal) pallidum and then onto the substantia nigra again, particularly to the pars reticulata (nonpigmented) cells, and an indirect system that traverses the lateral (external) pallidum and continues on to the subthalamic nucleus, with which it has strong to-and-fro connections In most ways, the subthalamic nucleus and lateral pallidum operate as a single functional unit (at least in terms of the effects of lesions in those locations on parkinsonian symptoms and the neurotransmitters involved; see further on) The medial pallidum and reticular part of the nigra can be viewed in a similar unitary way, sharing the same input and output patterns Within the indirect pathway, an internal loop is created by projections from the subthalamic nucleus to the medial segment of the pallidum and pars reticulata A second offshoot of the indirect pathway, recently uncovered, consists of projections from the lateral pallidum to the medial pallidonigral output nuclei These anatomic and physiologic concepts are continuously being revised on the basis of clinical and neurochemical observations As already mentioned, a complete account of this intricate connectivity cannot be given; but the main themes outlined here seem to have been accepted The relative importance of the direct and indirect pathways is being altered by recent experience, as discussed below From the internal pallidum, two bundles of bers reach the thalamus the ansa lenticularis and the fasciculus lenticularis The. how to make barcodes in excel free Barcode scanner software freeware chip - innachapoll.ml
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